Sabah - ID Request

[Zicky]

Hi Doctor Seow,

Had a amazing trip to Borneo, and would like to seek for some ID, the list is quite hefty, so I understand it will take some time, so no rush to reply.
I refer to the butterflies of borneo volume 2 written by the japanese as a reference:hmmm::thinking:

1 – Troides Helena?
inat link - https://www.inaturalist.org/observations/196345962

2 – Appias nero?
inat link - https://www.inaturalist.org/observations/196345941

3 - Saletara panda?
inat link - https://www.inaturalist.org/observations/196345935

4 – 7 – Chersonesia rahria?
4 – https://www.inaturalist.org/observations/196339550
5 - https://www.inaturalist.org/observations/196339574
6 - https://www.inaturalist.org/observations/196339619
7 - https://www.inaturalist.org/observations/196345960

8 – 10, Eurema hecabe?
8 - https://www.inaturalist.org/observations/196345960
9 - https://www.inaturalist.org/observations/196345903
10 - https://www.inaturalist.org/observations/196345913
11 – neptis haritia?
inat link - https://www.inaturalist.org/observations/196345924

12 – 14 - Symbrenthia lilaea?:thinking:
12 - https://www.inaturalist.org/observations/196339569
13 - https://www.inaturalist.org/observations/196345872
14 - https://www.inaturalist.org/observations/196345923

15 - Cirrochroa emalea?
inat link https://www.inaturalist.org/observations/196345922

16 – Pelopidas?
inat link - https://www.inaturalist.org/observations/196345915

17 – Tanaecia iapis?
inat link - https://www.inaturalist.org/observations/196345879

18 – Tirumala
inat link - https://www.inaturalist.org/observations/196339632

19 – 21 – Mycalesis pintana?
19 - https://www.inaturalist.org/observations/196339526
20 - https://www.inaturalist.org/observations/196345857
21 - https://www.inaturalist.org/observations/196345860

22 - Euploea camaralzeman?
inat link - https://www.inaturalist.org/observations/196339525

23 – celastrina?:confused:
inat link - https://www.inaturalist.org/observations/196331468

24 – 27 - Genus Notocrypta?
24 - https://www.inaturalist.org/observations/196345932
25 - https://www.inaturalist.org/observations/196339590
26 - https://www.inaturalist.org/observations/196331514
27 - https://www.inaturalist.org/observations/196339589

28 – 30 – Koruthaialos Sindu?:thinking:
28 - https://www.inaturalist.org/observations/196339594
29 - https://www.inaturalist.org/observations/196345900
30 - https://www.inaturalist.org/observations/196339539
31 - https://www.inaturalist.org/observations/196339545

32 – caltoris?
inat link - https://www.inaturalist.org/observations/196345871

33 – lambrix salasala?
inat link - https://www.inaturalist.org/observations/196345906

Skipping regards:gbounce::jumjoy::redbounce
Zicky

[Psyche]

https://www.inaturalist.org/observations/196345962
1. Female Troides helena mosychlus N. Borneo

Female T. helena mosychlus in Sabah.
https://www.flickr.com/photos/anschieber/49513092146

Female T. amphrysus flavicollis Sabah.
https://www.flickr.com/photos/berniedup/8067437684
https://a4.pbase.com/g10/25/686825/3/166051576.kFz9QoPO.jpg



https://www.inaturalist.org/observations/1963459412.
2. Male Appias nero.


https://www.inaturalist.org/observations/196345935
3. Male Saletara panda.

[Psyche]

https://www.inaturalist.org/observations/196339550
4. Chersonesia intermedia.

https://www.inaturalist.org/observations/196339574
5. Chersonesia rahria.

https://www.inaturalist.org/observations/196339619
6. Chersonesia rahria.

https://www.inaturalist.org/observations/196345960
7. Chersonesia rahria.


Species in which the FW 3rd band from the base is single (ie. not with double edgings.)

Fw submarginal band has wavy margin.

1a. FW apex crossed by a curved line/band........C. rahria.
https://4.bp.blogspot.com/-ZRXxVkCHCzM/U5BumcI7xKI/AAAAAAAAPDM/O1oMZCeSnuw/s1600/DSC_0353.JPG

1b. FW apex not crossed by a line/band.....C. intermedia.
https://wingscales.com/Nymphalidae/Chersonesia-intermedia-intermedia


2. FW submarginal band with straight margin.......C. risa
https://thaibutterflies.com/Butterflies/chersonesia-risa/

[Psyche]

https://www.inaturalist.org/observations/196345903

9. Uncertain. Too worn.


https://www.inaturalist.org/observations/196345913

10. Probably Eurema lacteola.

Two species can be very similar.
they both have a single FW cellspot. HW spot 7 is a sort of squiggle.

In E. andersoni the FW border in space 2 & 3 is roughly parallel to the margin.
https://blogger.googleusercontent.com/img/b/R29vZ2xl/AVvXsEhreKdVIxVNQQzwvHqASRO2U95uxvBV0eow-P3WefhjAnSZ12u7pFG_aS7MwrV59uMKUrwUviSTywDfQ9WDs8k tvH6EoTipTR3X-vECouwwF4YW_farQfR8HLWHd9SXPDr6xCbVOHPNs2w/s1600/EA_adult_Federick_01_c.jpg
https://blogger.googleusercontent.com/img/b/R29vZ2xl/AVvXsEhOGHSy_64viMj3HDQImJDLu7QJ-RJxBbcGhPEZu5H1-8s0krzGabKGgvXrBawHNSBs5Om5cc5eGfTx0jA4jzE0JlfdJwX XX1OTYUmFVszk6bTDaRBZifnTrOItz07MiHpyEZeAKFPQ2oA/s1600/EA_Khew_01.jpg

In E. lacteola, the fW border's inner margin in space 2 &3 is slanted from costa to tornus.
This alignment is similar to that of E. hecabe.
https://www.inaturalist.org/observations/112141147
https://www.inaturalist.org/observations/117280909

[Psyche]

https://www.inaturalist.org/observations/196345924

11. Female Neptis omeroda.

Three species may look similar.

Neptis harita.
This can be identified by the HW submarginal spots being catenulate,ie. like a chain-link.
male & female.
https://blogger.googleusercontent.com/img/b/R29vZ2xl/AVvXsEhse1DxYrE_TSwHvAja9lBf12I7LA56RFJF95XZVMoP9b e9xAXJQ99K3gt6RlllyBFt-bLGP0rKUWwgmHklAGgsZZdvfK8prnMvUZmhE0c6JEhLM0F6-yRCAgo0wxlagW31ssn1ewww0oU/s1600/CS_adult_newly_eclosed_02.jpg
https://www.butterflycircle.com/checklist/mugshots/Neptis%20harita%20harita/Chocolate%20Sailor%20-%20Bene%20Tay.jpg

Neptis omeroda.
This looks similar but the FW postdiscal spots are rounded (not concaved) & the Hw submarginal band is not catenulate.
https://yutaka.it-n.jp/lim1/720220001.html
https://thaibutterflies.com/wp-content/uploads/2016/07/Neptis-omeroda-1.jpg


Neptis ilira have rather dark narrow & obscure markings.
https://yutaka.it-n.jp/lim1/720240010.html

[Psyche]

https://www.inaturalist.org/observations/196339569
https://www.inaturalist.org/observations/196345872
https://www.inaturalist.org/observations/196345923
28502

12, 13 & 14.
Likely one species.
Should be Symbrenthia hippoclus.
The two species S. lilaea & hippoclus are very similar & a recent gene analysis suggest they may form a supercomplex.
This was based on a couple of gene markers & may be inadequate.
I follow Eliot in C&P4.

Symbrenthia hippoclus.
a Sundanian & Papuan species.
UnF discal spot 3 which is triangular is largely pinkish. Similar HW discal spots 4, 5, 6 may also be pinkish but hard to see in fieldshots.
UnH discal spot 7 more heavily shaded brown at the corner.
https://blogger.googleusercontent.com/img/a/AVvXsEjSvUbDzVDmmEJahPsCSHzsMLwm7b7CrSbCseYhsD8iJ4 wzic7pNchN98vwiH-aSX7-SmIttGJk2VNFI0vuEufhVGujyob6gDjmujGZyml6HFzpd-97pUOwQcENBotlll4puiW8cMCOXP0StDWepUfjQWUrlzQC9HTQ 9zxbLpwnpikjAzZb1Bz3j09n=s2000
https://blogger.googleusercontent.com/img/a/AVvXsEgxz9ruWNpzauBzSyjIsTWl_FWiLezOmjpwwVILxn6PNS 1u_g6aZGNBu7P9TTGBF0sDqdYu49Qx0CaekLRJogWIYniO_Ox-4U4shhoaTOIqqgA5zkDOrziqA97Kfk6t8h1_0-uGVlLT5oetjBLt6nxq9cIxp2PS_uFJSdoHTP8mt6_krfCFI-CjVU6e=s1600
https://blogger.googleusercontent.com/img/a/AVvXsEhsRttmTKpujDyyndCZMrIFyZRlknrZ9CIpMVQ7e4IB1L T2RUsoHGdb8aAcD4z1ugYbZ5DwwbeHLsGQqjtQ7F1B1m7t8VG1 _mYWqo17lzcbGrT3N1hRa3s-yHXaos0GD0xkXa5IbO9lHDBT2gCQjQyHJw2LWpKOIdzoKaq8-f_B9Vl3uus0HKdOIEvW=s1704
From Papua.
https://www.flickr.com/photos/angiud/45512555425


Symbrenthia lilaea.
UnF discal spot 3 tawny; UnH discal spot 7 margin largely clear; ground less dark-shaded.
https://thaibutterflies.com/Butterflies/simbrenthia-lilaea/

[Psyche]

https://www.inaturalist.org/observations/196345922

15. Cirrochroa malaya.

Two species are similar in which UpF have broad black border.

Cirrochroa emalea.
UpF apex with a pale spot in the black border.
UnH silvery band much constricted in the upper half.
https://www.butterflycircle.com/checklist/mugshots/Cirrochroa%20emalea%20emalea/MalayYeoman-KSK2.jpg
https://www.butterflycircle.com/checklist/mugshots/Cirrochroa%20emalea%20emalea/malay-yeoman-khew.jpg

Cirrochroa malaya.
UpF apex without a pale spot.
UnH silvery band only slightly narrower in the upper half.
https://www.flickr.com/photos/chris_chafer/28622156893
https://www.inaturalist.org/observations/161033946

[Psyche]

https://www.inaturalist.org/observations/196345915

16. Female Isma cronus.

Female Isma is notoriously difficult to ID, but this one is clear enough.

unH with a minute white spot in the cell & in space 2.
Underside of abdomen banded.

See C&P4 page 366 key line 8.



https://www.inaturalist.org/observations/196345879

17. Male Tanaecia cocytina ambalika.
Previously as Tanaecia iapis ambalika.
https://inaturalist.nz/observations/69258956
https://inaturalist.nz/observations/65339509#data_quality_assessment


The nominate subspecies in Java ie. T iapis iapis is very different, being marked in white.
https://www.jungledragon.com/image/107238/horsfields_baron_java_tanaecia_iapis_iapis.html/zoom
https://www.jungledragon.com/image/99961/horsfields_baron_java_tanaecia_iapis_iapis.html/zoom

Most authors now relegate the forms with blue band in the male as T. cocytina including on the continent.
The form in Singapore would be T. cocytina puseda.
it is also placed in the genus Cynitia by some.
https://yutaka.it-n.jp/lim2/721070010.html



https://www.inaturalist.org/observations/196339632

18. Male Tirumala septentrionis miosticta.
The forms from Borneo, including those of I]Ideopsis[/I] have very narrow markings.
Continental form. Note male dark brand at bases of space 2 & 3.
https://www.ifoundbutterflies.org/sites/default/files/558_73863-666-5faaf007b3cbe-1.jpg

[Psyche]

https://www.inaturalist.org/observations/196339526
https://www.inaturalist.org/observations/196345857
https://www.inaturalist.org/observations/196345860

19, 20, & 21 all Mycalesis anapita.

M. pitana.
This Bornean endemic is IDed thus.
Upperside with broad black border on both wing; FW ocelli totally obliterated.
UnF with an eyespot in space 3 ,so no gap between the spots.
Underside dark discal fascia with a branch marking the cellend, less clear on the FW.
https://static.inaturalist.org/photos/104013920/large.jpg
https://inaturalist-open-data.s3.amazonaws.com/photos/245641505/large.jpeg
https://commons.wikimedia.org/wiki/Category:Mycalesis_pitana#/media/File:Mycalesis_pitana_(25004347283).jpg


The other two can be confusing especially on the underside.
UnF without a spot in space 3, so there is a gap between spot 2, & spots 4 & 5.


Mycalesis anapita
UpF with the FW eyespot clear of the black border or at least 3/4 clear.
UnF with upper third of the brown discal fascia not darker.
unF with the upper submarginal spot 5 usually much larger than spot 4.
Ssp. fucienta from Borneo have clearer ocellus.
https://www.jamiun.com/wp-content/uploads/2023/07/208A9954watermarksmall.jpg
https://www.jamiun.com/wp-content/uploads/2023/07/208A9980watermarksmall.jpg
https://www.jamiun.com/wp-content/uploads/2018/02/IMG_4818watermarksmall.jpg


Mycalesis patiana.
UpF with the ocellus in space 2 largely merged into the black border.
UnF dark discal fascia blackish in the upper third ,often after a kink.
UnF submarginal spots 4 & 5 sometimes more equal in size.
P. malaysia.
https://2.bp.blogspot.com/-1QV0YDqhkSw/V0-Ej1EcjfI/AAAAAAAAaBs/FG63TpiOCw44hX_mmaGmc-5ZYXqxrN3YQCLcB/s1600/DSC_2410.JPG
https://nlliew66butterflies.blogspot.com/2016/09/mycalesis-patiana-eliot.html
Sabah.
https://uk.inaturalist.org/observations/24634638
https://www.flickr.com/photos/dave_irving/9828106165/in/photostream/

[Psyche]

https://www.inaturalist.org/observations/196339525

22. Euploea camaralzeman scudderi.
The FW is marked by 3-4 series of small white spots.

The Bornean subspecies have much reduced spotting on the HW.
https://1.bp.blogspot.com/-BXTkI8B-oSM/Vgbul-5u9DI/AAAAAAAAUh4/HzaYp0VoiVA/s400/Malayan%2BCrow%2B%2528Euploea%2Bcamaralzeman%2Bscu dderi%2529%2BDSC_7582.jpg




https://www.inaturalist.org/observations/19633146823.
23. Should be Celastrina lavendularis placidina.

The markings are as for Celastrina.
Three species are known in Borneo.

In C. philippina & algernoni, the UnH submarginal line defining the black spots is rounded (crenate).
In C. lavendularis placidina the line is sharply serrate.
Also C. lavendularis tend to be darker grey with white-ringed spots.
See all three below.
http://tropical-lycaenidae.net/tropical-lycaenidae/lycaenopsis/celastrina/

[Psyche]

https://www.inaturalist.org/observations/196345932
24. Female Notocrypta paralysos.
FW band broad & not reaching the costa full-width. FW with spot 4.

https://www.inaturalist.org/observations/196339590
25. Notocrypta pria.
FW band narrow & tapers towards the costa & dorsum (lower margin).
https://yutaka.it-n.jp/hes/91590001.html


https://www.inaturalist.org/observations/196331514
26. Notocrypta paralysos.
FW band broad, not reaching the costa, its inner margin with a right-angled notch; FW with a spot 4.


https://www.inaturalist.org/observations/196339589
27. Very likely female Notocrypta paralysos variant.
FW with a large spot 4 & a tiny spot 5.
FW band broad, with a right angled notch on its inner margin.
Band reaches costa, but yellowish .


Notocrypta paralysos
https://yutaka.it-n.jp/hes/91600010.html
FW band broad with a distinct right-angled notch; FW spot 4 usually present.
UnF bant not reaching costa or only patchily.
FW shape more rounded.
Males.
https://blogger.googleusercontent.com/img/b/R29vZ2xl/AVvXsEhSxgtaqmQdzfzl9DTXOhyAYNykfcChp0V-e-6wXKdC498gzbPIERF_D06hv2bAfeR3Htebw9VW6_QfKE1bEtu3 AbV6ebc-4qg_PUTcDGAXANuBB2aZWk0cjzPuwXfXBMW_qgc0UP5wfTBg/s1600/BD_adult_Jonathan.JPG
https://www.jamiun.com/wp-content/uploads/2017/10/IMG_3670watermarksmall.jpg
Females.
https://blogger.googleusercontent.com/img/b/R29vZ2xl/AVvXsEhwx6nAVKoBR-fDvd_Iu_QaLquqt7iCWcE8wiKr4LGBoz5CR4Vf3rRRf6sroDa5 K6TDvReIbft_Mt-twumxVt3m6oQPvpRAVzE_tsbmL2RphbmVpL54MNgzmHy-7_HBcZITjlGqOH6cJCm-/s1600/BD_newly_eclosed_underside_lateral_01.jpg
https://blogger.googleusercontent.com/img/b/R29vZ2xl/AVvXsEio_QCcJSdtMeAOJuksq9s4BDm-IaMudvkrZITNtQwA5A-FFe8y8WWhRJJYFlHrDf9r5gxCoDEKdM8P-EKkM7yMA_GDr2nGaksoa_Q8DKQ08HPDWL1A6ROIE1MuMyNPYTE kNuo1rG3D2Hez/s1600/BD_adult_EC_01.jpg


Notocrypta clavata.
https://yutaka.it-n.jp/hes/91610001.html
FW band narrower with notch on the inner margin more obtuse.
FW spot 4 rarely present.
FW sharper.
UnF band broadly extend to the costa.
Males.
https://www.flickr.com/photos/gancw1/13027025594/
https://commons.wikimedia.org/wiki/Category:Notocrypta_clavata#/media/File:Notocrypta_clavata_theba_(Evans,1949)_(156234 33394).jpg
Probable female.
https://thaibutterflies.com/wp-content/uploads/2016/07/Notocrypta-clavata.jpg

[Psyche]

https://www.inaturalist.org/taxa/475716-Koruthaialos
28. Koruthaialos rubecula female.
Orange band not reaching costa.

https://www.inaturalist.org/observations/196345900
29. Korithaialos sindu.
Orange band reaching costa.


https://www.inaturalist.org/observations/196339539
30. Probably Koruthaialos frena.
Similar but larger than K. sindu.
FW band very broad reaching costa, its outer border no mid-margin notch.
Band 's lower part reaching termen (outer margin) broadly.


https://www.inaturalist.org/observations/196339545
31. Probably K. sindu. band reaches costal margin.


K. rubecula.
FW orange generally narrow, & not quite reaching costa. Female with broader band.
Outer margin without a notch.
3rd segment of palpi long & narrow.
https://yutaka.it-n.jp/hes/91510001.html
https://pamsbutterflies.com/butterfly/1475/koruthaialos-rubecula


K. sindu.
FW orange band reaching costa. with outer mid-margin notch.
Palpi 3rd segment short & conical.
https://yutaka.it-n.jp/hes/91530001.html
https://4.bp.blogspot.com/-6RkAaBJ000s/T-K3aDr6a_I/AAAAAAAAGNA/OxkxMDv66sE/s640/DSC1400++Koruthaialos+sindu.jpg


K. frena
Larger than K. sindu. Hill insect.
FW band uniformly very broad, reaching costa & the lower outer margin(termen) to the tornus.
Orange band's outer border without a mid-margin notch.
Palpi 3rd segment short.


Ancistroides (Kerana ) gemmifer.
UnH marked with silvery spots.
https://yutaka.it-n.jp/hes/91580001.html
https://nl.wikipedia.org/wiki/Bestand:Gem_Demon_(Ancistroides_g._gemmifer)_(1672 5854595).jpg


Ancistroides 9Kerana) armatus.
Large FW 23-27mm, Impressive.
FW orange baand with a distinct upperr notch.
https://yutaka.it-n.jp/hes/91570001.html



https://www.inaturalist.org/observations/196345871
32. Probably a worn out Caltoris cormasa.


https://www.inaturalist.org/observations/196345906
33. Iambrix stellifer.
The position of the two HW spots in pic 2 match that of I. stellifer.
https://yutaka.it-n.jp/hes/91480001.html
https://yutaka.it-n.jp/hes/91470001.html


TL Seow: CHeers.

[Zicky]

Thanks Doctor Seow for the comprehensive explanation and the drawings. Will take the time to look through and read up. Appreciate it